High resolution macro of Wasp found in Greece
Wenzel and mantis
Tiger swallowtail, Papilio glauca, and Platanthera dilatata orchid


Native Wildflower Gardens

To serve a growing interest in gardening with native plants, I have tried over the years to demonstrate certain species that do well in the central Appalachian area, along with a few traditional non-native favorites of special note. Effort was made to exhibit plants that differ from common sun-loving garden plants, such as those that favor shade or can inhabit a rain garden. A rain garden serves to capture water runoff and put it back in the soil, and it is populated by plants that can tolerate both inundation (when it rains) and drought (between rains.) A sandy garden represents plants that like well-drained soils. The attached list of nearly 200 species is divided first by habitat type (sunny, shady, etc.) then by estimated bloom period.  This list may provide inspiration and helpful information for aspiring gardeners. Most of these plants are implemented in the Virtual Garden Timeline.

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Drone models

Unmanned Aerial Vehicles (drones) have become a basic tool, enabling us to explore and document our surroundings as never before. Powdermill scientists took the lead in developing drone-based photogrammetry, a method that combines hundreds or thousands of photos taken from different angles, to create a 3D model of the forest at the level of an entire stand. A trick we used was to eliminate the crowns so that we can more accurately model the tree trunks (which is why they look like they are dead, no twigs!) A model of one of our research plots was composed of 23 million reference points, and the dimensions of the trees is correct to within about 3 cm. You can vist the model here:

Virtual Garden Timeline

The woodland wildflowers of eastern North America are beautiful and diverse. Many of them bloom so early that there are no leaves on the trees and people are not yet thinking of getting out in the woods. With a team that included a botanist, a horticulturist, and several digital artists, we created an animated summary of what flowers bloom when in the area near Powdermill Nature Reserve. Certain of these plants are available as Virtual Reality models that can be navigated on your computer screen. An award-winning phone app allows you to plant these flowers wherever you point your phone!

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Everyone wants clean water, whether for our own use or as part of a healthy natural environment. Water quality of streams is generally determined by monitoring the insects that live in the stream, with healthier streams producing a greater variety of specialist insects. As part of a team of entomologists, computer programmers, and learning researchers, we created an extraordinary teaching resource that turns your computer screen into a microscope that you drive. Expert instructions show how to identify insects important in water quality analyses. A phone app permits some of the same utility in a portable fashion.

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YearAuthorsTitlePrint PublicationWeb publication
2023Spicer, M.E., A.A. Royo, J.W. Wenzel, and W.P. CarsonUnderstory plant growth forms respond independently to combined natural and anthropogenic disturbances.Forest Ecology and Management 543(3): 121077https://doi.org/10.1016/j.foreco.2023.121077
2022Swartz, K.A., T.C. Weathers, N.M. Zembower, D. Wu, A. Kautz, B. Stouffer, J.W. Wenzel, and J.E. Carlson Genetic Variation and Conservation Implications in Pennsylvania for the Rare and Imperiled Buffalo Nut (Cervantesiaceae; Pyrularia pubera Michaux).Annals of Carnegie Museum 87 (3), 235-248
2022Kautz, A.R., M.R. Louw, C. Bartley, J.C. Morse, and J.W. WenzelMacroinvertebrates.org: Creating a Digital Teaching Collection of Common Freshwater Macroinvertebrates of Eastern North AmericaAmerican Entomologist 68: 28–31https://doi.org/10.1093/ae/tmac012
2021DeGroote, L.W., E. Hingst-Zaher, L. Moreira-Lima, J.V. Whitacre, J.B. Slyder, and J.W.WenzelCitizen science data reveals the cryptic migration of the Common Potoo Nyctibius griseus in Brazil.Ibis 163:380-389
2021Noll, F.B., M. da Silva, R.A. Soleman, R.B. Lopes, Y.C. Grandinete, E.A.B. Almeida, J.W. Wenzel, and J.M. Carpenter.Marimbondos: systematics, biogeography, and evolution of social behavior of neotropical swarm-founding wasps (Hymenoptera: Vespidae:Epiponini). Cladistics 37: 423-441.
2021Perry, K.I., F.S. Sivakoff, K.F. Wallin, J.W. Wenzel, and D.A. Herms.Forest disturbance and arthropods: Small-scale canopy and understory disturbances alter movement of mobile arthropods,Ecosphere 12(11):e03771.10.1002/ecs2.3771
2020Rangel, J., B. Traver, M. Stoner, A. Hatter, B. Trevelline, C. Garza, T. Shepherd, T.D. Seeley, and J. Wenzel. Genetic diversity of wild and managed honey bees (Apis mellifera) in Southwestern Pennsylvania, and prevalence of the microsporidian gut pathogens Nosema ceranae and N. apis. Apidologie. Doi:10.1007/s13592-020-00762-5.
2020Wenzel, J. W. Nest structure: Social wasps.14pp, In: Encyclopedia of Social Insects, C. Starr, (ed.) doi.org/10.1007/978-3-319-90306-4_146-1
2020Wenzel, J. W.Le SRAS et le COVID-19 n’ont pas de lien avec les civettes et les pangolins. (SARS and COVID-19 do not come from civets and pangolins).Bulletin de la Société Française de Systématique 62: 14-22.
2020Wenzel, J. W. Origins of SARS-CoV-1 and SARS-CoV-2 are often poorly explored in leading publications.Cladistics 36: 374-379.doi.org/10.1111/cla.12425
2019Harrington, M. C. R., M. Tatzgern, T. Langer, and J. W Wenzel.Augmented reality brings the real world into natural history dioramas with data visualizations and bioacoustics at the Carnegie Museum of Natural History.Curator: The Museum Journal 62:177-193.
2019Slyder, J.B., J. W. Wenzel, A. A. Royo, M. E. Spicer, W. P. Carson. Post-windthrow salvage logging increases seedling and understory diversity with little impact on composition immediately after logging.New Forests, doi.org/10.1007/s11056-019-09740-x
2018Curtze, A.C., T.A. Carlo, and J.W. Wenzel.The effects of a tornado disturbance and a salvage timber extraction on the seed-rain and recruitment community of an eastern temperate deciduous forest.Northeast Naturalist, 25: 627-645.
2018Harrington, Maria C. R., J. W. Wenzel, M. Tatzgern, T. Langer, M. Oliver, B. Isaac, A. Guffey, Z. Bledsoe, C. Jones, R. Dinic, R., and M. Tiefengrabner. The AR Perpetual Garden Apps.Apple iTunes App Store, or Google Play App Store, developer: The Carnegie Institute, URL available on demand.
2018Krishnadas, M., N.G. Beckman, J.C. Zuluaga, Y. Zhu, J. Whitacre, J.W. Wenzel, S.A. Queenborough, and L.S.Comita. Environment and past land use together predict functional diversity in a temperate forest.Ecological Applications 28.8: 2142-2152.DOI: 10.1002/eap.1802
2018Perry, K.I., K.F. Wallin, J.W. Wenzel, and D.A. Herms. Forest disturbance and arthropods: Small-scale canopy gaps drive invertebrate community structure and composition.Ecosphere 9: e02463, doi: 10.1002/ecs2.2463
2018Spicer, M.E., K. F. Seuss, J. W. Wenzel, and W. P. CarsonDoes salvage logging erase a key physical legacy of a tornado blowdown: A case study of tree tip-up mounds?Canadian Journal of Forest Research 48: 976-982.Dx.doi.org/10.1139/cjfr-2018-0037
2018Wenzel, J. W. Dr. Eleanor’s Book of Common Ants. [Book review].American Entomologist 64: 125
2018Wickert, K.L, A. M. Metheny, D. D. Davis, D. M. Geiser, J. W. Wenzel, D. Planinsek, and M. T. Kasson. First report of Fusarium stem canker on Pyrularia pubera, a rare native parasitic shrub in forests of southwestern Pennsylvania. Plant Disease 102: 1852.
2017Perry, K. I., K. F. Wallin, J. W. Wenzel, and D. A. Herms.Characterizing movement of ground-dwelling arthropods with a novel mark-capture method using fluorescent powder.J. Insect Behavior 30: 32-47, DOI 10.1007/s10905-017-9598-0
2017Fotis, A., S. Murphy, R. Ricart, M. Krishnadas, J. Whitacre, J. Wenzel, S. Queenborough, and L. Comita.Aboveground biomass is driven by mass-ratio effects and stand structural attributes in a temperate deciduous forest.J. Ecology 106 (2), 561-570.
2017Angulo, E, G. M. Luque, S. D. Gregory, J. W. Wenzel, C. Bessa?Gomes, L. Berec, and F. Courchamp.Allee effects in social species.J. Animal Ecology 87: 47-58
2017Harrington, Maria C. R., Wenzel, J. W., Oliver, M., Isaac, B., Guffey, A., Bledsoe, Z. and C. Jones.The Virtual Garden Timeline Websitehttps://virtualgarden.powdermill.org/
2016Andena, S. R, M. Aragão, and J. W. Wenzel.As vespas sociais encontradas no Semiárido brasileiro. [Social wasps found in the Brazilian semiarid region].Pp 126-138 in Conhecendo Os Artrópodes Do Semiárido, [Understanding the arthropods of the semiarid]. F. Bravo and A.R. Calor, Eds., Métis Produção Editorial, São Paulo.
2015Murphy, S.J., L.D. Audino, J. Whitacre, J.L. Eck, J.W. Wenzel, S. Queenborough, and L.S. Comita. Species associations structured by environment, space, and land use history promote beta-diversity within a relatively homogeneous temperate forest. Ecology 96: 705-715.
2015Slyder, J. J. Whitacre, and J Wenzel. Painting a clearer picture of shale gas development.ArcNews Spring 2015: 20-21.
2015Calinger, K.M, E.A. Calhoon, H. Chang, J. Whitacre, J. Wenzel, L.S. Comita, and S.A. Queenborough. Historic Mining and Agriculture as Indicators of Occurrence and Abundance of Widespread Invasive Plant Species.PLOS ONE June 5, 2015; DOI: 10.1371/journal.pone.0128161
2015Zeigler D., and J. W. Wenzel.Paläste wie aus Pappmaché [Palaces of paper maché].Pages 134-137 in Wissens Dinge: Geschichten aus dem naturkundemuseum. [Knowledge of things: Stories from the natural history museum] Nicolai Press. Berlin
2014Randle, C.P., and J. W. Wenzel.Research Note: Deer browsing threatens a locally rare parasitic plant, buffalo nut (Pyrularia pubera Michx.)J. Pennsylvania Academy of Science 88: 231-234.
2013Carpenter, J.M., S.R. Andena, F. B. Noll, and J. W. Wenzel.Well, what about intraspecific variation? Taxonomic and phylogenetic characters in the genus Synoeca de Saussure (Hymenoptera, Vespidae)?Zootaxa 3682: 421-431.
2012Whitacre, J. and J. W. Wenzel.Carnegie Museum of Natural History Pennsylvania Unconventional Gas Well Data.Published on the web, URL available on demand.
2010Wenzel, J. W.Les charactèrs comportementaux sont de très bons indicateurs de phylogenie. (Behavioral characters are good indicators of phylogeny)Biosystema (Paris) 27: 3-18.
2010Wenzel, J. W.Stepping into the light: review of "Biological Systematics - Principles and Applications" second edition by Randall T. Schuh and Andrew V. Z. Brower. [Book review].BioEssays 32:260-262.
2010Oliveira, O. A. L. de, F. B. Noll, and J. W. Wenzel.Foraging behavior and colony cycle of Agelaia vicina (Hymenoptera: Vespidae; Epiponini).J. Hymenoptera Research 19: 4-11.
2010Mynhardt, G. and J. W. Wenzel. Phylogenetic analysis of the myrmecophilous Cremastocheilus Knoch (Coleoptera, Scarabaeidae, Cetoniinae), based on external adult morphology. Zookeys 24: 129-140.
2010Noll, F. B., B Gomes, A.C.O. Lima, S. Mateus and J. W. Wenzel.Castes in the neotropical social wasp Leipomeles dorsata (Fabricius)(Hymenoptera: Vespidae): a window for workers achieving a new status in the colony.Neotropical Entomology 39: 549-554.
2010Bucheli, S.R., S. Passoa, and J. W. WenzelA phylogenetic test of Ehrlich and Raven’s theory of escape and radiation in insects that feed on toxic plants, based on Nearctic Depressaria moths (Gelechioidea: Elachistidae: Depressariinae), with a discussion of the evolution of genitalia. Entomologica Americana 116: 1-24.
2009Blackledge T.A., N. Scharff, J. Coddington, T. Szüts, J.Wenzel, C. Hayashi, I. Agnarsson.Reconstructing web evolution and spider diversification in the molecular era.Proceedings of the National Academy of Science (USA): 106: 5229-5234.
2009Mynhardt, G, and J. W. Wenzel.Voyages of Discovery: Exploration of Plants and Animals Through Space and Time.32pp. Museum of Biological Diversity, Ohio State University.
2009Wenzel, J. W., and P. DeWitt.The Struggles and the Glory of the Pre-Darwinians.52pp. Museum of Biological Diversity, Ohio State University.
2009Felippotti, G.T., G. M. Tanaka Junior, F. B. Noll, and J. W. Wenzel.Discrete dimorphism among castes of the bald faced hornet Dolichovespula maculata (Hymenoptera: Vespidae) in different phases of the colony cycle. J. Natural History, 43: 2481-2490.
2009Caesar, R. M., and J. W. Wenzel.A Phylogenetic Test of Classical Species Groups in Argia (Odonata: Coenagrionidae).Entomologica Americana 115: 97-108.
2008Noll, F. B., and J. W. Wenzel. Caste in the swarming wasps: "Queenless" societies in highly social insects. Biological Journal of the Linnean Society 93: 509-522.
2008Wenzel, J. W. Alien Invaders: Introduced Species And Their Environmental Impact.32pp. Museum of Biological Diversity, Ohio State University.
2008Song, H., and J. W. Wenzel. Mosaic pattern of genital divergence in three populations of Schistocerca lineata Scudder, 1899 (Orthoptera: Acrididae; Cyrtacanthacridinae). Biological Journal of the Linnean Society 94: 289-301.
2008Song, H., and J. W. Wenzel. Phylogeny of bird-grasshopper subfamily Cyrtacanthacridinae (Orthoptera: Acrididae) and the evolution of locust phase polyphenism.Cladistics 24: 515- 542.
2008Wenzel, J. W., and G. M. Luque.Parsimony cluster analysis in successional ecology.Cladistics 24: 746-757.
2008Gilligan, T., and J. W. Wenzel.Extreme intraspecific variation in Hystrichophora genitalia ­Questioning the lock-and-key hypothesis (Lepidoptera: Tortricidae).Annales Zoologici Fennici 45: 465-477.
2008Wenzel, J. W. Built by Animals: The Natural History of Animal Architecture. [Book review].Quarterly Review of Biology 83: 320-321.
2007Pickett, K. M., and J. W. Wenzel.Revision and cladistic analysis of the nocturnal wasp genus Apoica Lepeletier (Hymenoptera: Vespidae; Polistinae, Epiponini). American Museum Novitates #3562, 1-30.
2007Raczkowski, J. R. and J. W. Wenzel.Biodiversity studies and their foundation in taxonomic scholarship.Bio Science 57: 974-979.
2006Bucheli, S.R., D. Horn, and J. W. Wenzel. Biodiversity of Gelechioidea (microlepidoptera): Sampling to assess a re-established Appalachian forest in Ohio based on gelechioid moths (Lepidoptera: Gelechioidea).Biodiversity and Conservation 15: 503-516.
2006Naug, D., and J. Wenzel.Constraints on foraging success due to resource ecology limit colony productivity in social insects.Behavioral Ecology and Sociobiology 60: 62-68.
2006Abdul-Salim, K., M. Daly, J.V. Freudenstein and J.W. Wenzel.Report on the Ohio State University workshops in phylogenetic methods. [Editorial].Cladistics 22: 387-389.
2006Wenzel, J.W. and F.B. Noll. Dados comportamentais na era da genômica [Behavioral data in the era of genomics].Etologia 8: 63-69.
2005Pickett, K. M., S. R. Rissing, and J. W. Wenzel.Iconoclasts of Evolution: Haeckel, Behe and Wells and the ontogeny of a fraud.American Biology Teacher 67:270-277.
2005Bucheli, S.R., and J. W. WenzelGelechioidea (Insecta: Lepidoptera) systematics: A reexamination using combined morphology and mitochondrial DNA data.Molecular Phylogenetics and Evolution 35: 380-394.
2005Pickett, K. M., G. L. Tolman, W.C. Wheeler, and J. W. Wenzel.Parsimony overcomes statistical inconsistency with the addition of more data from the same gene.Cladistics (21) 438-445 .
2004Wenzel, J. W. Why I teach introductory courses.Pages 21-28 In: Talking About Teaching. Ohio State University Academy of Teaching.
2004Wenzel, J.W., K. C. Nixon, and G. Cuccodoro.Dites non au Phylocode!Bulletin de la Societé Française de Systématique 31: 19-23.
2004Pickett, K. M. and J. W. WenzelPhylogenetic analysis of the New World Polistes (Hymenoptera: Vespidae) using morphology and molecules.J. Kansas Entomol. Soc. 77: 742-­760.
2004Noll, F. B., J. W. Wenzel, and R. Zucchi.Evolution of caste in Neotropical swarm-founding wasps (Hymenoptera: Vespidae; Epiponini).American Museum Novitates #3467, 1-24.
2003Branham, M.A., R.A. Broadley, and J.W. Wenzel. 2003Bioluminescence in insects.Pp. 33-36 In: Yearbook of Science and Technology, 2003. McGraw-Hill Publishers, New York.
2003Branham, M.A. and J.W. Wenzel. The Origin of Photic Behavior and the Evolution of Sexual Communication in Fireflies (Coleoptera: Elateroidea).Cladistics 19: 1-22.
2003Freudenstein, J. V., K. M. Pickett, M. P. Simmons, and J.W. Wenzel. From basepairs to birdsongs: phenotypic data in the age of genomics.Cladisitcs 19: 333-348.
2003Wenzel, J. W. A user's guide to the tree of life. [Book Review].Cladistics 19: 376-378.
2002Simmons, M.P., C.P. Randle, J.V. Freudenstein, and J.W. Wenzel. Limitations of relative apparent synapomorphy analysis (RASA) for measuring phylogenetic signal.Molecular Biology and Evolution 19(1):14-23.
2002Wenzel, J. W. The role of the Willi Hennig Society in Systematics [Editorial].Cladistics 18: 1-3.
2002Bucheli, S.R., J. -F. Landry, and J. W. Wenzel.Larval case architecture and implications for host plant associations in North American Coleophora (Lepidoptera: Coleophoridae).Cladistics 18: 71-93.
2002Wenzel, J. W. Phylogenetic analysis: The basic method.Pages 4- 30, in R. DeSalle, G. Giribet, and W.Wheeler, eds., Techniques in Molecular Systematics, Birkhäuser, Basel.
2001Blackledge, T.A. and J.W. Wenzel. State-determined foraging decisions and web architecture in the spider Dictyna volucripes (Araneae, Dictynidae). Ethology Ecology Evolution 13: 105-113.
2001Blackledge, T.A. and J.W. Wenzel. Silk mediated defense in an orb-web spider against predatory mud dauber wasps.Behaviour. 138: 155-172.
2001Pickett, K.M., Osborne, D.M., Wahl, D., and Wenzel, J.W. 2001. An enormous nest of Vespula squamosa from Florida, the largest social wasp nest reported from North America, with notes on colony cycle and reproduction.  J. New York Entomol. Soc. 109: 408-415.
2001Branham, M.A. and J.W. Wenzel. The Evolution of Bioluminescence in Canthariods (Coleoptera: Elateroidea).Florida Ent. 84: 565-586.
2000Karsai, I., and J. W. Wenzel.Organization and regulation of nest construction behavior in Metapolybia wasps.Journal of Insect Behavior: 13: 111-140.
2000Carpenter, J. M., J. -I. Kojima and J. W. Wenzel.Polybia, paraphyly, and polistine phylogeny.
2000Blackledge, T. A, and J. W. Wenzel. The evolution of cryptic spider silk: A behavioral test.Behavioural Ecology 11: 142-145.
2000Pickett, K. M, A. McHenry, and J. W. Wenzel.Nestmate recognition in the absence of a pheromone.Insectes Sociaux 47: 212-219.
2000Pickett, K. M, and J. W. Wenzel. High productivity in haplometrotic colonies of the introduced paper wasp Polistes dominulus (Hymenoptera: Vespidae; Polistinae).J. New York Entomol. Soc.: 108: 314-325.
1999Wenzel, J. W., and M. E. Siddall.Noise.Cladistics 14: 51-64.
1999Blackledge, T. A, and J. W. Wenzel.Do stabilimenta in orb webs attract prey or defend spiders?Behavioural Ecology 10: 372-376.
1999Carpenter, J. M., and J. W. Wenzel.The relative abundance of swarm-founding social wasps in the Congo Basin (Insecta: Hymenoptera; Vespidae, Polistinae).Natural History Bulletin of Ibaraki University 3: 9-14.
1998Wenzel, J. W. A generic key to the nests of hornets, yellowjackets and paper wasps world-wide (Vespidae: Vespinae, Polistinae).American Museum Novitates #3224, 1-39.
1998Karsai, I., and J. W. Wenzel.Individual- and colony-level flexibility, productivity, and organization of work as consequences of colony size.Proceedings of the National Academy of Science (USA) 95: 8665-8669.
1998Kojima, J., J. W. Wenzel, J. M. Carpenter.The types of Icaria galimatia de Saussure 1853 and I. ignobilis de Saussure, 1990, with establishment of their synonymy (Hymenoptera: Vespidae: Polistinae). Entomological Science 1: 393-401.
1997Wenzel, J. W. When is a phylogenetic test good enough?Memoires du Museum National D’Histoire Naturelle (Paris) 173: 31-45.
1996Wenzel, J. W. Learning, behaviour programs, and higher-level rules in nest construction of Polistes.Pages 58-74. In S. Turillazzi and M. J. West-Eberhard, eds., Natural History and Evolution of Paper Wasps. Oxford Press, Oxford.
1996Carpenter, J. M., J. W. Wenzel, & J. Kojima.Synonymy of the genus Occipitalia Richards, 1978, with Clypearia de Saussure 1854 (Hymenoptera: Vespidae; Polistinae, Epiponini). Journal of Hymenoptera Research 5: 157-165.
1996Karsai, I., Z Pénzes, & J. W. Wenzel.Dynamics of colony development in Polistes dominulus: a modeling approach.Behavioral Ecology and Sociobiology 39: 97-105.
1995Miller, J. S., & J. W. Wenzel.Ecological characters and phylogenetic inference in the study of insects.Annual Review of Entomology 40: 389-415.
1995Karsai, I., & J. W. Wenzel.Nests built on the dorsum of conspecifics in Polistes: the value of anomolous behavior.Animal Behaviour 50: 1429-1431.
1995Farris, J. S., M. Kallersjo, V. A. Albert, M. Allard, A. Anderberg, B. Bowditch, C. Bult, J. M. Carpenter, T. M. Crowe, J. De Laet, K Fitzhugh, D. Frost, P. Goloboff, C. J. Humphries, U. Jondelius, D.Judd, P. O. Karis, D. Lipscomb, M. Luckow, D. Mindell, J. Muona, K. Nixon, W. Presch, O. Seberg, M. E. Siddall, L. Struwe, A. Tehler, J. Wenzel, Q. Wheeler and W. Wheeler.Explanation.Cladistics 11: 211-218.
1994Wenzel, J. W., & J. M. Carpenter.Comparing methods: Adaptive traits and tests of adaptationPages 79-101. In P. Eggleton and R. Vane-Wright, eds., Phylogenetics in Ecology. Harcourt Brace & Co., London.
1993Wenzel, J. W. Application of the biogenetic law to behavioral ontogeny: a test using nest architecture in paper wasps.J. Evolutionary Biology 6: 229-247.
1993Stubblefield, J. W., J. Seger, J. W. Wenzel, & M. M. Heisler. Temporal, spatial, sex-ratio and body-size heterogeneity of prey species taken by the beewolf Philanthus sanbornii (Hymenoptera: Sphecidae).Philosophical Transactions of the Royal Society of London. B 339: 397-423.
1992Wenzel, J. W. Extreme queen-worker dimorphism in Ropalidia ignobilis, a small-colony wasp (Hymenoptera:Vespidae).Insectes Sociaux 38: 31-43.
1992Wenzel, J. W. What do you get if you cross a cladist with a behavioral ecologist? [Book review].Cladistics 8: 97-102.
1992Wenzel, J. W. Les nids de guêpes sociale d'interêt historique conservés au MNHN Paris. [The social wasp nests of historical interest preserved in the MNHN, Paris.] Revue Française d'Entomologie (N.S.) 14: 1-11.
1992Wenzel, J. W. Behavioral homology and phylogeny.Annual Review of Ecology and Systematics 23: 261-281.
1991Wenzel, J.W. The evolution of nest architecture in the social vespids.Pages 480-519. In K. G. Ross and R. W. Matthews, eds., Social Biology of Wasps, Cornell Univ. Press, Ithaca.
1991Wenzel, J. W., & J. Pickering.Cooperative foraging, productivity, and the central limit theorem.Proceedings of the National Academy of Science (USA) 88: 36-38.
1990Wenzel, J. W. A social wasp's nest from the Cretaceous period, Utah, USA, and its biogeographical significance.Psyche 97: 21-29.
1990Espelie, K. E. , J. W. Wenzel, & G. Chang. Surface lipids of the social wasp Polistes metricus Say and of its nest and nest pedicel and their relation to nestmate recognition.J. Chemical Ecology 16: 2229-2241.
1989Wenzel, J. W. Endogenous factors, external cues and eccentric construction in Polistes annularis (Hymenoptera: Vespidae).J. Insect Behavior 2: 679-699.
1989Carpenter, J. M. & J. W. Wenzel. Synonymy of the genera Protopolybia and Psuedochartergus (Hymenoptera: Vespidae, Polistinae).Psyche 96: 177-186.
1988Smith, B. H. & J. W. Wenzel.Pheromonal covariation and kinship in social bee Lasioglossum zephyrum Hymenoptera: Halictidae).J. Chemical Ecology 14: 87-94.
1988Carpenter, J. M. & J. W. Wenzel.A new species and nest type of Mischocyttarus from Costa Rica, with descriptions of nests of three related species.Psyche 95: 89-99.
1987Wenzel, J.Male reproductive behavior and mandibular glands in Polistes major (Hymenoptera: Vespidae).Insectes Sociaux 34: 44-57.
1987Wenzel, J.Ropalidia formosa, a nearly solitary paper wasp from Madagascar (Hymenoptera: Vespidae).J. Kansas Entomological Society 60: 549-556.